Many groups around the world continue to study Rubisco activase with the ultimate goal of determining whether alterations will be able to improve the photosynthetic efficiency of plants. Ogren’s remarkable mentorship PF-02341066 cell line : The Lifetime Achievement Award also recognizes that in addition to his own extraordinary research accomplishments, Ogren has provided outstanding leadership as a mentor and leaves a scientific legacy that includes a remarkable progression of students and postdoctoral associates. Less well known outside the UIUC campus is the fact that he was instrumental in several highly successful USDA and University

of Illinois at Urbana-Champaign faculty hires. Several of these students, postdocs and faculty have become world leaders in their own right.

One of the more compelling, but lesser-known examples of his excellence in recognizing and promoting young talent is that he successfully nominated one of his graduate students, Jeff Werneke, for a quadrennial award in 1989 from the Council of Graduate Schools for the Distinguished Dissertation in Biological Sciences. Jack Widholm We end this News Report of the Ceremony where Ogren was honored with a testimonial by Jack Widholm; Jack continues to work at the UIUC, and has known Bill Ogren for more than 40 years. The Widholm and Ogren families are close friends. Jack wrote: It is a great honor for me to be a part of the Ogren Lifetime Achievement Award Ceremony. We have done work together and been friends since 1968. I am not a photosynthesis person but in 1967 when I was working at the International Minerals and Chemical CX-4945 datasheet Corporation in MM-102 in vivo Libertyville, Illinois I had an idea about how to screen for plants that lacked

photorespiration. The idea was to grow C3 plants under low CO2 conditions below the CO2 compensation concentration where they would lose CO2 and die. I wrote a letter to the USDA to get funding, I got none, but the letter made it to Bill in USDA and he responded that it might be a good idea. Interestingly in May 1968, Dichloromethane dehalogenase I joined the Agronomy Department at the University of Illinois at Urbana-Champaign (UIUC), and, thus, Bill and I worked together on the idea (Widholm and Ogren 1969). We showed that indeed C3 but not C4 plants would die under low CO2; we then screened the oat collection and about 350,000 mutagenized soybean plants with no survivors! (For a historical perspective on C-3 pathway, see Benson 2005; and Bassham 2005; and for C-4 pathway, see Hatch 2005.) Clearly, if we had succeeded in eliminating photorespiration, the yields of many crops would have increased greatly, but we did not, and later work by Bill Ogren and Chris Somerville with Arabidopsis showed that the photorespiratory pathway cannot be blocked and still have viable plants. Thus attempts to alter Rubisco to not react with oxygen have not yet been successful.